Keep me in the loop, I'm keen to know the difference too. Seeing as they are miles apart phylogenetically, there should be some morphological differences in the pupa that are easily observed in the exuviae.
Where the pupa was found is useful info, but not definitive. Some info from Common: Endoxyla encalypti, the most common large cossid in our part of Australia, feeds internally at the base of the trunk and into the roots of Acacia. It pupates underground and the pupa protrudes from the ground before emergence. (Other Endoxyla species have other foodplants, eg E. liturata uses Eucalyptus). Abantiades hepialids feed on the roots of Eucalyptus from the outside and similarly pupate underground and the pupa protrudes from the ground before emergence. So a pupa found nearby to the base of an Acacia is a good prospect for E. encalypti, and one found further out from the trunk of a eucalypt is a prospect for an Abantiades. Where was this pupa originally?
It was found in an area (a nature strip actually) that I cleared of small native vegetation late last year and subsequently covered with 'forest waste' wood chip mulch. The mulch had been in place since well before the fires hit nearby on NewYears day and I walk the space regularly checking for weeds so the shell had not been there for more than a day or so. There is a large long dead tree stump and miscellaneous other trees in the area. The moth must be enormous!
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